All but one scientist was pivotal in making some type of discovery that led to our understanding of the basic nature of cells. that scientist is

Hans Krebs as the scientist who was pivotal in making some type of discovery that led to our understanding of the basic nature of cells.

Hans Krebs (25 August 1900 – 22 November 1981) was a  British physician and biochemist born in Germany. He discovered that nutrients are broken down in our cells to release energy for the construction of cells

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during the process of dna replication, brdu can replace thymidine and incorporate into the newly synthesized dna, cell proliferation thus can be reflected in the extent of brdu incorporation.