They are a layered structure located between paired homologs in prophase i of meiosis in many organisms. they are required for moving homologs to opposite sides of the poles during anaphase i of meiosis. they are required for moving sister chromatids to opposite sides of the poles during anaphase ii of meiosis and anaphase of mitosis. they catalyze the phosphorylation of specific proteins. they are important for maintaining homologous chromosome pairing and plays a role in recombination. they are filaments in mitotic and meiotic spindles.